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Were the demographic changes a direct result of the selectivity of the epidemic, i. Or were improvements in diet and other standards of living more important causal factors in these demographic trends?

These two possibilities are not necessarily mutually exclusive, as both were ultimately the results of the massive mortality caused by the Black Death.

Nonetheless, the disentanglement of intrinsic versus extrinsic factors that could have led to changes in survival and mortality is relevant to an understanding, more generally, of the effects of disease on human biology and social, political, and economic conditions.

Such a disentanglement, however, requires further study, such as an analysis of stable isotopes and skeletal stress markers that are associated with nutritional deficiencies before and after the Black Death to determine if diet changed in substantial ways that are discernible from the skeleton.

Analysis of nitrogen isotope values, for example, might reveal whether people in general consumed substantially more animal protein following the Black Death than was true before the epidemic [66].

The results of this study also raise questions about the possible effects of migration, i. The improvements in mortality and survivorship observed in the post-Black Death sample might indicate that migration into the city after the epidemic introduced a large number of healthy people.

According to Dyer [29] migration likely increased after Black Death as an expression of resistance against restrictions enacted under labor laws in England, such as attempts to prevent increases in wages after However, London drew substantial numbers of migrants from throughout England and beyond throughout the medieval period, both before and after the Black Death [67] , [68].

Famine, in particular, spurred migration before the Black Death, and London attracted large numbers of rural residents in search of work and charity [67].

The existence of migration throughout the medieval period means that both the pre-Black Death and post-Black Death assemblages likely contained a mixture of immigrants and native Londoners, so differences between the two cannot be attributed only to the effects of migration.

Furthermore, the possibility of increased immigration to London following the Black Death does not explain the differences in age-at-death distributions observed between the two time periods, the most striking of which is the higher proportion of older adults in the post-Black Death sample.

The differences in the age-at-death distributions would reflect the effects of migration alone only if the majority of migrants were older adults; however, it is more likely that most migrants were late teens or young adults [32] , [69].

Lastly, the Black Death spared few regions within Europe [70]. Therefore, the selective mortality that operated during the epidemic would have shaped health and demography throughout the continent.

Migrants into London in the post-Black Death period would have included survivors of the epidemic and their descendants and the health and demographic characteristics thereof to the same or similar extent as did the native population of London.

In summary, the existence of migration into London following the Black Death does not necessarily undermine the conclusions made here about positive changes in mortality and, by inference, health following and resulting from the Black Death, thought it is certainly an important issue worthy of further study.

The results of this study indicate that mortality and survivorship improved in the generations following the Black Death, and that the patterns observed are not simply an artifact of temporal changes in fertility.

These results highlight the power that infectious diseases have to shape population-wide patterns of health and demography over both the short- and long-term.

Site codes and contexts for all individuals included in the study. I am grateful to Jelena Bekvalac and Rebecca Redfern at the Museum of London Centre for Human Bioarchaeology for providing access to the skeletal samples used in this study and for generously providing the physical facilities for this work.

I also thank Dr. Eric Jones, Dr. Andrew Noymer, and three anonymous reviewers for providing comments on this manuscript. Conceived and designed the experiments: SND.

Performed the experiments: SND. Analyzed the data: SND. Wrote the paper: SND. Browse Subject Areas? Click through the PLOS taxonomy to find articles in your field.

Abstract The medieval Black Death c. Introduction The Black Death was one of the most devastating epidemics in human history. Materials and Methods 2.

Pre-Black Death Samples: St. Post-Black Death Sample: St. Mary Graces. Download: PPT. Figure 1. Age-at-death distributions for the pre- and post-Black Death samples.

Figure 2. Kaplan-Meier survivorship functions for the pre- and post-Black Death samples. Table 1. Maximum likelihood estimates with standard errors of the Gompertz parameters and the effect of the time period covariate on mortality with likelihood ratio test.

Table 2. Discussion The age-at-death distributions from the pre- and post-Black Death samples suggest that survival improved following the Black Death, as the post-Black Death sample has a higher proportion of older adults.

Conclusions The results of this study indicate that mortality and survivorship improved in the generations following the Black Death, and that the patterns observed are not simply an artifact of temporal changes in fertility.

Supporting Information. Table S1. Acknowledgments I am grateful to Jelena Bekvalac and Rebecca Redfern at the Museum of London Centre for Human Bioarchaeology for providing access to the skeletal samples used in this study and for generously providing the physical facilities for this work.

References 1. Cambridge: Cambridge University Press. London: Arnold. Human Biology — View Article Google Scholar 4. Twigg G The Black Death: a biological reappraisal.

London: Batsford Academic and Educational. Hinde A England's population: a history since the domesday survey. J Archaeol Sci — View Article Google Scholar 7.

View Article Google Scholar 8. View Article Google Scholar 9. Usher BM A multistate model of health and mortality for paleodemography: Tirup cemetery.

Demography — View Article Google Scholar Med Hist Suppl 74— Comp Stud Soc Hist 8: — Nottingham Medieval Studies — Nature — Emerging Infectious Diseases — J Infect Dis — Evolution — Econ Hist Rev — Bailey M T.

Ashton Prize: Joint Winning Essay. Econ Hist Rev 1— Dyer C Making a living in the middle ages: the people of Britain Hatcher J Plague, population, and the English economy, — London: Macmillan.

Rappaport SL Worlds within worlds: structures of life in sixteenth-century London. Stone DJ The consumption of field crops in late medieval England.

Food in medieval England: diet and nutrition. Oxford Oxford University Press. Econ Hist Rev 2: — Dyer C An Age of Transition?

Scrimshaw NS Historical concepts of interactions, synergism and antagonism between nutrition and infection. J Nutr S—S. Past Present 33— Paine RR If a population crashes in prehistory, and there is no paleodemographer there to hear it, does it make a sound?

Am J Phys Anthropol — London: London and Middlesex Archaeological Society. Mary Spital. London: Museum of London Archaeology.

Schofield J Excavations on the site of St. Transactions of the London and Middlesex Archaeological Society 77— The London Archaeologist 5: — Museum of London Archaeology Service Monograph London: Museum of London Archaeology Service.

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In the summer of , Silent Circle announced that they would be releasing a successor to the Blackphone, the Blackphone 2, in September Blackphone 2 does not have a removable battery.

Kismet Smart Wi-Fi Manager comes pre-installed. It also includes an international power adapter kit and a headset. Ars Technica praised that the Blackphone's Security Center in PrivatOS gives control over app permissions, such as the bundled Silent Phone and Silent Text services that anonymise and encrypt communications so no one can eavesdrop on voice, video, and text calls.

They also praised the Disconnect VPN and Search that keeps web trackers from the telephone and anonymises web searches and Internet traffic.

The telephone's original launch quantity is unknown, but was reported to have sold out shortly after the launch began.

As of June [update] , a Blackphone was on exhibit at the Victoria and Albert Museum and one had been added to the collection of the International Spy Museum.

In , Silent Circle had significant financial problems, caused by a significant overestimate of how many phones they could sell.

This led to the near bankruptcy of the company. From Wikipedia, the free encyclopedia. For the phone by Boeing, see Boeing Black. This article needs additional citations for verification.

Please help improve this article by adding citations to reliable sources.

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